top banana UK /ˌtɒp bəˈnɑːnə/US /ˌtɑp bəˈnænə/ noun2 •(informal) The boss, the leader.Only the top banana can make a decision of that magnitude. •(informal, old-fashioned) The principal comedian in a vaudeville or burlesque show.He was hilarious, and remained the top banana on the tour for years. formstop bananas