epibranchial adjective 1noun 1forms 1 adjective1 •Of or pertaining to a segment, below (more ventral than) the pharyngobranchial segment and above (more dorsal than) the ceratobranchial segment, on either side of a branchial arch (in a fish) or of a pharyngeal arch (in the embryo of a vertebrate).According to T. J. Parker, they probably represent the epibranchial region of the second branchial arch, and Beddard also refers them to the same arch. — 1920 noun1 •An epibranchial cartilage or bone.The fourth and fifth epibranchials of other genera have the same tendency as these clupeids and salmonids. — 1969 formsepibranchials