nucleobranch adjective 1noun 1forms 1 adjective1 •(no longer used) Belonging to the Nucleobranchiata, a obsolete order of molluscs that have the gills packed in the shell along with the heart, such sea snails now mostly classified in the order Littorinimorpha. noun1 •(no longer used) One of the order Nucleobranchiata. formsnucleobranchs